Josh Hough
In Memory of Josh Hough, PhD (Department of Ecology & Evolutionary Biology 2016) 

Josh was a highly accomplished graduate student who was passionate about addressing fundamental questions in evolutionary biology. Josh’s interests in evolutionary genetics brought him to the University of Toronto, where he pursued a PhD under the supervision of Spencer Barrett and Stephen Wright.

Sadly, Josh passed away in British Columbia on Sunday, September 16, 2018. He will be dearly missed by his family and his many colleagues and friends, who will always remember his independence, fearlessness, creativity, and passion for knowledge.